Beach Spawning
Redd excavation
With her alpha male guarding behind her, a female Sockeye Salmon (Oncorhynchus nerka) excavates her redd during preparation to beach spawn in Eyak Lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiEcologyInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfemale behaviorfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismredd excavationsemelparitysemelparousunder fresh waterunderwater
Beach spawning act
Sockeye Salmon (Oncorhynchus nerka) in the act of beach spawning in Eyak Lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiEcologyInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismsemelparitysemelparousspawningspawning actspawning behaviorunder fresh waterunderwater
Spawning commencement
Sockeye Salmon (Oncorhynchus nerka) commencing to beach spawn in Eyak lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismsemelparitysemelparousspawningspawning actspawning behaviorunder fresh waterunderwater
Covering redd
Sockeye Salmon (Oncorhynchus nerka) pair following beach spawning in Eyak lake, Alaska
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiEcologyInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipcovering redddiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismsemelparitysemelparousspawning behaviorunder fresh waterunderwater
Gametes mingling
Eggs and sperm mingle as Sockeye Salmon (Oncorhynchus nerka) beach spawn in Eyak lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiEcologyInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismsemelparitysemelparousspawningspawning actspawning behaviorunder fresh waterunderwater
Beach spawning act
Sockeye salmon (Oncorhynchus nerka) commencing to beach spawn in Eyak Lake, Alaska. Eggs being extruded into redd pocket. Shoreline is to the left (note alge on rocks visible through male's gape.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiEcologyInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismredsemelparitysemelparousspawningspawning actspawning behaviorunder fresh waterunderwater
Beach spawning act
Sockeye salmon (Oncorhynchus nerka) commencing to beach spawn in Eyak Lake, Alaska. First eggs being extruded into redd pocket. Shoreline is to the left (note alge on rocks visible through male's gape.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiEcologyInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismredsemelparitysemelparousspawningspawning actspawning behaviorunder fresh waterunderwater
Beach spawning act
Sockeye salmon (Oncorhynchus nerka) commencing to beach spawn in Eyak Lake, Alaska. Female making final anal fin probe into spawning pocket before extruding eggs. Shoreline is to the left (note alge on rocks visible through male's gape).
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiEcologyInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismredsemelparitysemelparousspawningspawning actspawning behaviorunder fresh waterunderwater
Redd excavation
Sockeye salmon (Oncorhynchus nerka) redd excavation prior to beach spawning in Eyak Lake, Alaska. Female excavates the redd by making rapid body undulations. Shoreline is to the left.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfemale behaviorfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismredd excavationsemelparitysemelparousunder fresh waterunderwater
Redd excavation
Sockeye salmon (Oncorhynchus nerka) redd excavation prior to beach spawning in Eyak Lake, Alaska. Female excavates the redd by making rapid body undulations. Shoreline is to the left.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalchordatecourtshipdiadromousdiadromyfemale behaviorfresh waterfreshwaterlakelenticlivingmigratorynear Cordovaorganismredd excavationsemelparitysemelparousunder fresh waterunderwater
Probing
As her alpha male comes alongside, a female Sockeye Salmon (Oncorhynchus nerka) uses her anal fin to probe her redd prior to beach spawning in Eyak Lake, Alaska. The shoreline is to the left in this image.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismprobingsalmon habitatsemelparitysemelparousspawning behaviorunder fresh waterunderwaterwildwild salmon
Rejection
A female Sockeye Salmon (Oncorhynhus nerka) attacks an unwanted intruding male while she is in the process of preparing her redd to beach spawn in Eyak Lake, Alaska. Her favored alpha male is in the background.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateaggressionanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparousunder fresh waterunderwaterwildwild salmon
Rejection
Female Sockeye Salmon (Oncorhynchus nerka) attempts to nudge away an intruding male while preparing her redd to beach spawn in Eyak Lake, Alaska
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateaggressionanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparousunder fresh waterunderwaterwildwild salmon
Circular swimming
Female Sockeye salmon circles back towards her redd during its excavation in preparation for beach spawning in Eyak Lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparousunder fresh waterunderwaterwildwild salmon
Staging Sockeye Salmon
Male Sockeye Salmon (Oncorhynchus nerka) staging in beach spawning habitat in Eyak lake, Alaska
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparousstagingunder fresh waterunderwaterwildwild salmon
Redd excavation
A partially spawned-out female Sockeye Salmon (Oncorhynchus nerka) excavates her redd while beach spawning in Eyak lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfemale behaviorfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismredd excavationsalmon habitatsemelparitysemelparousunder fresh waterunderwaterwildwild salmon
The gathering
Sockeye Salmon (Oncorhynchus nerka) staging prior to beach spawning near the mouth of a small creek in Eyak Lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparousstagingunder fresh waterunderwaterwildwild salmon
The gathering
Sockeye Salmon (Oncorhynchus nerka) staging prior to beach spawning near the mouth of a small creek in Eyak Lake, Alaska.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparousstagingunder fresh waterunderwaterwildwild salmon
Beach spawning topside
Topside view of Sockeye salmon (Oncorhynchus nerka) beach spawning site in Eyak Lake, Alaska. Absence of algae indicative of a redd excavation activity or wave action (at shoreline).
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparoustopsideunder fresh waterwildwild salmon
Beach spawning topside
Topside view of Sockeye salmon (Oncorhynchus nerka) beach spawning site in Eyak lake, Alaska with camera set up. Note speed limit sign indicating road trhat follows lake shoreline.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorRunning Bear CreekSalmonidaeSalmoniformesSalmoninaeSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratorynaturenear Cordovaorganismsalmon habitatsemelparitysemelparoustopsideunder fresh waterunderwater techniquewildwild salmon
Beach spawning topside
Sockeye Salmon (Oncorhynchus nerka) on beach spawning redds along the shore of Eyak Lake, Alaska near an unamed creek. Redd depressions appear dark rather than the glacial green of offshore waters seen in the background due to seepage of groundwater that makes this site conducive to beach spawning.
AlaskaBiologyEcologyReproductive behaviorSouth Central AlaskaSummerUnited Statesanadromousanadromybeach spawningbiologicalcourtshipdiadromousdiadromyfresh waterfreshwaterlakelenticlivingmigratoryorganismsalmon habitatsemelparitysemelparoustopsideunder fresh water
Beach spawning Sockeye Salmon
Sockeye Salmon a.k.a. Red Salmon (Oncorhynchus nerka) preparing to beach spawn in Eyak Lake, Alaska during the summer.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCordovaEcologyEyak LakeInfraclass TeleosteiNeopterygiiOncorhynchus nerkaPacific salmonPhylum ChordataPrince William SoundRed SalmonReproductive behaviorSalmonidaeSalmoniformesSalmoninaeSeacam underwater housingSnell's windowSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterin situlakelenticlivingmigratorynaturalworldnatureorganismsalmon habitatsemelparitysemelparousunder fresh waterunderwaterunderwaterphotowildwild salmon
Beach spawning
Topside view of Sockeye Salmon (Oncorhynchus nerka) beach spawning in Eyak Lake, Alaska during the summer. Polecam used to take underwater photographs is seen.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCoho SalmonCordovaEcologyEyak LakeHumpback SalmonInfraclass TeleosteiNeopterygiiOncorhynchus gorbuschaOncorhynchus kisutchOncorhynchus nerkaPacific salmonPhylum ChordataPink SalmonPrince William SoundRed SalmonReproductive behaviorSalmonidaeSalmoniformesSalmoninaeSalvelinus malmaSalvelinus malma lordiSeacam underwater housingSilver SalmonSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterin situlakelenticlivingmigratorynaturalworldnatureorganismprobingsalmon habitatsemelparitysemelparousspawning behaviorunder fresh waterunderwaterunderwater techniqueunderwaterphotowildwild salmon
Beach spawning
Topside view of Sockeye Salmon (Oncorhynchus nerka) beach spawning in Eyak Lake, Alaska during the summer. Polecam used to take underwater photographs is seen.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCoho SalmonCordovaEcologyEyak LakeHumpback SalmonInfraclass TeleosteiNeopterygiiOncorhynchus gorbuschaOncorhynchus kisutchOncorhynchus nerkaPacific salmonPhylum ChordataPink SalmonPrince William SoundRed SalmonReproductive behaviorSalmonidaeSalmoniformesSalmoninaeSalvelinus malmaSalvelinus malma lordiSeacam underwater housingSilver SalmonSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterin situlakelenticlivingmigratorynaturalworldnatureorganismprobingsalmon habitatsemelparitysemelparousspawning behaviorunder fresh waterunderwaterunderwater techniqueunderwaterphotowildwild salmon
Beach spawning
Sockeye Salmon (Oncorhynchus nerka) beach spawning in Eyak Lake, Alaska during the summer.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCoho SalmonCordovaEcologyEyak LakeHumpback SalmonInfraclass TeleosteiNeopterygiiOncorhynchus gorbuschaOncorhynchus kisutchOncorhynchus nerkaPacific salmonPhylum ChordataPink SalmonPrince William SoundRed SalmonReproductive behaviorSalmonidaeSalmoniformesSalmoninaeSalvelinus malmaSalvelinus malma lordiSeacam underwater housingSilver SalmonSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterin situlakelenticlivingmigratorynaturalworldnatureorganismprobingsalmon habitatsemelparitysemelparousspawning behaviorunder fresh waterunderwaterunderwaterphotowildwild salmon
Beach spawning
Sockeye Salmon (Oncorhynchus nerka) beach spawning in Eyak Lake, Alaska during the summer.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCoho SalmonCordovaEcologyEyak LakeHumpback SalmonInfraclass TeleosteiNeopterygiiOncorhynchus gorbuschaOncorhynchus kisutchOncorhynchus nerkaPacific salmonPhylum ChordataPink SalmonPrince William SoundRed SalmonReproductive behaviorSalmonidaeSalmoniformesSalmoninaeSalvelinus malmaSalvelinus malma lordiSeacam underwater housingSilver SalmonSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterin situlakelenticlivingmigratorynaturalworldnatureorganismprobingsalmon habitatsemelparitysemelparousspawning behaviorunder fresh waterunderwaterunderwaterphotowildwild salmon
Beach spawning
Sockeye Salmon (Oncorhynchus nerka) beach spawning in Eyak Lake, Alaska during the summer.
ActinopterygiiAlaskaAnimalBiologyChordataClass ActinopterygiiCoho SalmonCordovaEcologyEyak LakeHumpback SalmonInfraclass TeleosteiNeopterygiiOncorhynchus gorbuschaOncorhynchus kisutchOncorhynchus nerkaPacific salmonPhylum ChordataPink SalmonPrince William SoundRed SalmonReproductive behaviorSalmonidaeSalmoniformesSalmoninaeSalvelinus malmaSalvelinus malma lordiSeacam underwater housingSilver SalmonSockeye SalmonSouth Central AlaskaSubclass NeopterygiiSummerTeleostTeleosteiUnited StatesVertebrateanadromousanadromybeach spawningbiologicalbiospherechordatecourtshipdiadromousdiadromyfresh waterfreshwaterin situlakelenticlivingmigratorynaturalworldnatureorganismprobingsalmon habitatsemelparitysemelparousspawning behaviorunder fresh waterunderwaterunderwaterphotowildwild salmon