Schooling

Salmonid fishes will aggregate into schools when migrating and when holding in deeper areas such as pools in streams.

Coho Salmon spawning migration
A male Coho Salmon (Oncorhynchus kisutch) begins to asert himself by using the gaping threat while schooled in a deep pool during the last stages of maturation in the Autumn in an Alaska, USA stream.
Actinopterygii Alaska Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Power Creek Reproductive behavior Salmonidae Salmoniformes Salmoninae Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate aggression anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater gaping in situ living lotic migratory naturalworld near Cordova organism schooling semelparity semelparous spawning migration staging stream under fresh water underwater
Putative stray Chinook Salmon
Topside view of the odd stray Chinook Salmon (Oncorhynchus tshawytscha) among Pink Salmon (O. gorbuscha) and Chum Salmon (O. keta) in a dense school homing to spawn in Hartney Creek near Cordova, Alaska during summer. The small size suggests that this stray is a jack salmon
Actinopterygii Alaska Alaska wild salmon Animal Biology Chinook Salmon Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Hartney Creek Humpback Salmon Infraclass Teleostei King Salmon Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Oncorhynchus tshawytscha Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate biological biosphere chordate courtship covering redd in situ jack jack salmon living naturalworld nature near Cordova organism photographed under water precocial male precocious male schooling spawning behavior straying topside underwater underwater photography underwaterphoto wild wild salmon
Chum Salmon female probing
With her mouth slightly agape and body flexed downward, a female Chum Salmon (Oncorhynchus keta) probes the gravely bottom near the edge of a stream (bottom of frame) in preparation for spawning while another female guards just downstream alongside many other salmon schooled in deeper water in this topside view.
Alaska PacificSalmon probing SalmonBehavior Actinopterygii Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate biological biosphere chordate courtship covering redd in situ living naturalworld nature organism photographed under water spawning behavior topside underwater underwater photography underwaterphoto wild wild salmon
Recent arrivals from the ocean
Recent arrivals from the ocean enroute to spawning photographed just above the intertidal in a Prince William Sound, Alaska tributary stream. A male Pink Salmon (Oncorhynchus gorbuscha) asserting himself by gaping while schooled alongside conspecifics and Chum Salmon (O. keta). Photographed under water using a Canon EOS-1DX camera in a Seacam underwater housing equipped with an 8-15mm fisheye lens set at 15mm and a Seacam remote control.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Hartney Creek Infraclass Teleostei Neopterygii Oncorhynchus keta Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Snell's window South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate biological biosphere chordate courtship covering redd in situ living naturalworld nature near Cordova organism photographed under water schooling spawning behavior underwater underwater photography underwaterphoto wild wild salmon
Coho Salmon undergoing final maturation
Coho Salmon (Oncorhynchus kisutch) undergoing final maturation in a deep pool near their spawning grounds
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate diadromous diadromy fresh water freshwater glacial sediment in situ living lotic migratory naturalworld nature organism semelparity semelparous staging stream under fresh water underwater underwaterphoto wild wild salmon
Autumnal cohorts
While waiting for a spawning partner this sexually mature male Coho Salmon (Oncorhyncus kisutch) lurks in a pool occupied by resting Dolly Varden Charr (Salvelinus malma) during their autumnal migration to winter habitat in an Alaskan stream.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Dolly Varden Dolly Varden Char Dolly Varden Charr Dolly Varden Charr southern form Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Salmonidae Salmoniformes Salmoninae Salvelinus malma Salvelinus malma lordi Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate WPY2022 biological biosphere chordate in situ living naturalworld nature organism underwater underwaterphoto wild wild char wild charr wild salmon www.salmonography.com
Chum Salmon staging near their spawning grounds
Chum Salmon (Oncorhynchus keta) staging near their spawning grounds
Actinopterygii Alaska Alaska wild salmon Animal Biology Chinook Salmon Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Infraclass Teleostei King Salmon Neopterygii Oncorhynchus keta Oncorhynchus tshawytscha Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water semelparity semelparous spawning behavior staging stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Staging Sockeye, Chum, and Coho salmon
Sockeye (Oncorhynchus nerka), Chum (O. keta), and Coho (O. kisutch) salmon staging on their spawning migration.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Coho Salmon Dog Salmon Ecology Fish Infraclass Teleostei Neopterygii Oncorhynchus keta Oncorhynchus kisutch Oncorhynchus nerka Pacific salmon Palmer Hay Flats Sate Game Refuge Phylum Chordata Rabbit Slough Red Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon Sockeye Salmon South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate biological biosphere chordate courtship covering redd in situ living naturalworld nature organism photographed under water spawning behavior staging underwater underwater photography underwaterphoto wild wild salmon
Coho Salmon staging near their spawning grounds
Coho Salmon (Oncorhynchus kisutch) staging near their spawning grounds
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate WPY2022 biological biosphere chordate in situ living naturalworld nature organism staging underwater underwaterphoto wild wild salmon
Heads or tails?
Mature male Sockeye Salmon (Oncorhynchus nerka) swimis into a near ocean-bright immature Coho Salmon (O. kisutch) while looking for a mate in early autumn in an Alaskan stream.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Oncorhynchus nerka Pacific salmon Phylum Chordata Red Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon Sockeye Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate WPY2022 anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism run timing semelparity semelparous stream under fresh water underwater wild wild salmon
Early and late salmon arriving at spawning grounds
Nearing the end of their summer spawning period, mature Sockeye and Pink salmon (Oncorhynchus nerka and O. gorbuscha) school near their spawning grounds in early autumn coinciding with the arrival of numerous of Coho Salmon (O. kisutch) that commence spawning at the start of autumn.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Oncorhynchus nerka Pacific salmon Phylum Chordata Red Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon Sockeye Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature nuptial coloration ocean bright organism run timing semelparity semelparous staging stream under fresh water underwater underwaterphoto wild wild salmon
Sockeye and Coho salmon
Sockeye and Coho salmon (Oncorhynchus nerka and kisutch) nearing the end of their spawning migration schooled in a pool prior to spawning.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate biological biosphere chordate courtship in situ living naturalworld nature nuptial coloration organism run timing underwater underwaterphoto wild
The one that got away
Coho Salmon with Pixie lure. Aspect ratio = 2:3.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Fishing Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate diadromous diadromy fishing lure fresh water freshwater in situ living lotic migratory naturalworld nature organism semelparity semelparous stream under fresh water underwater underwaterphoto wild wild salmon
Coho Salmon spawning migration
Ocean bright Coho Salmon (Oncorhynchus kisutch) on their spawning migration in an Alaska USA stream at the start of Autumn. Although they are in fresh water their bright coloration suggests they have to wait some time before becoming fully mature and able to spawn.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Power Creek Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature near Cordova ocean bright organism semelparity semelparous stream under fresh water underwater underwaterphoto wild wild salmon
Coho Salmon spawning migration
Coho Salmon (Oncorhynchus kisutch) migrating to their spawning grounds.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism run timing semelparity semelparous stream under fresh water underwater underwaterphoto wild wild salmon
Sockeye Salmon emerging from glacial water
Sockeye Salmon (Oncorhynchus nerka) emerging from glacial water in a staging area. Gaping male signaling his dominance is at the lead. Aspect Ratio = 1:1.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Ecology Infraclass Teleostei Neopterygii Oncorhynchus nerka Pacific salmon Phylum Chordata Red Salmon Salmonidae Salmoniformes Salmoninae Seacam underwater housing Sockeye Salmon South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism semelparity semelparous staging stream under fresh water underwater underwaterphoto wild wild salmon
Sockeye Salmon spawning migration
Sockeye Salmon (Oncorhynchus nerka) spawning migration. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Copper River Delta Ecology Infraclass Teleostei Neopterygii Oncorhynchus nerka Pacific salmon Phylum Chordata Red Salmon Reproductive behavior Saddlebag Creek Salmonidae Salmoniformes Salmoninae Seacam underwater housing Sockeye Salmon South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ jack jack salmon living lotic migratory naturalworld nature nuptial coloration ocean bright organism precocial male precocious male run timing scale resorption semelparity semelparous staging stream under fresh water underwater underwaterphoto wild wild salmon
Sockeye Salmon spawning migration
Sockeye Salmon (Oncorhynchus nerka) staging during their spawning migration. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Copper River Delta Ecology Infraclass Teleostei Neopterygii Oncorhynchus nerka Pacific salmon Phylum Chordata Red Salmon Saddlebag Creek Salmonidae Salmoniformes Salmoninae Seacam underwater housing Sockeye Salmon South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism semelparity semelparous staging stream under fresh water underwater underwaterphoto wild wild salmon
Sharing the stream
Dolly Varden (Salvelinus malma) migrating upstream to spawning grounds alongside Pink and Chum salmon (Oncorhynchus gorbuscha and O. keta). Aspect Ratio = 1:2.5.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate WPY2022 biological biosphere chordate courtship covering redd diadromous fresh water freshwater in situ iteroparity iteroparous living lotic migratory naturalworld nature organism photographed under water schooling spawning behavior stream under fresh water underwater underwater photography underwaterphoto wild wild char wild charr wild salmon wild trout
Chum and Pink salmon spawning migration
Chum and Pink salmon (Oncorhynchus keta and O. gorbuscha) staging on their spawning migration. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Hartney Creek Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate WPY2022 aggression anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater gaping in situ living lotic migratory naturalworld nature near Cordova organism photographed under water semelparity semelparous spawning behavior spawning migration staging stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Maturing Coho Salmon
Still bright maturing Coho Salmon (Oncorhynchus kisutch) holding inear a spawning area. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Oncorhynchus nerka Pacific salmon Phylum Chordata Red Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon Sockeye Salmon South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism semelparity semelparous spawning migration stream under fresh water underwater underwaterphoto wild wild salmon
Chinook Salmon prior to spawning
Chinook Salmon (Oncorhynchus tshawytscha) prior to spawning in tannin-stained stream. Aspect ratio = 2:3.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chinook Salmon Chordata Class Actinopterygii Ecology Infraclass Teleostei King Salmon Neopterygii Oncorhynchus tshawytscha Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Snell's window South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water semelparity semelparous spawning migration stream under fresh water underwater underwaterphoto wild wild salmon
Chinook Salmon holding prior to spawning
Chinook Salmon a.k.a. King Salmon (Oncorhynchus tshawytscha) schooled in a holding pattern prior to spawning in a tributary of the Deshka River, Alaska, USA during the summer.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chinook Salmon Chordata Class Actinopterygii Deshka River tributary Ecology Infraclass Teleostei King Salmon Moose Creek Neopterygii Oncorhynchus tshawytscha Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Snell's window South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water salmonography.com semelparity semelparous spawning migration stream under fresh water underwater underwaterphoto wild wild salmon
Coastal Cutthroat Trout
Coastal Cutthroat Trout (Oncorhynchus clarkii clarkii) gather downstream of a beaver dam in a Copper River Delta, Alaska, USA stream during the spring.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Coastal Cutthroat Trout Copper River Delta Ecology Infraclass Teleostei Neopterygii Oncorhynchus clarkii Oncorhynchus clarkii clarkii Pacific trout Phylum Chordata Pipeline Lake Four Creek Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Spring Subclass Neopterygii Teleost Teleostei United States Vertebrate biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ iteroparity iteroparous living lotic migratory naturalworld nature organism salmonography.com searun spawning migration stream under fresh water underwater underwaterphoto wild wild trout
Maturing Coho Salmon
Coho Salmon (Oncorhynchus keta) at various stages of final maturation. Aspect Ratio = 2:3.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism semelparity semelparous spawning migration stream under fresh water underwater underwaterphoto wild wild salmon
Chumpy: A putative Chum and Pink salmon hybrid
A putative female Chum and Pink (Oncorhynchus keta and O. gorbuscha) salmon hybrid migrating upstream alongside mostly Pink Salmon. This Chumpy has spots, a color pattern, and teeth that are intermediate between the two parent species. Underwater view in an Alaskan stream during the summer.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Ecology Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater hybrid in situ living lotic migratory naturalworld nature organism semelparity semelparous spawning migration stream under fresh water underwater underwaterphoto wild wild salmon
Pink salmon spawning migration
Potential alpha male Pink Salmon (Oncorhynchus gorbuscha) gaping during spawning migration. Underwater view in an Alaskan stream during the summer.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate aggression anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic male behavior male threat posture migratory naturalworld nature organism photographed under water salmonography.com schooling semelparity semelparous spawning behavior spawning migration stream threat posture under fresh water underwater underwater photography underwaterphoto wild wild salmon
Pink Salmon spawning migration
Pink Salmon (Oncorhynchus gorbuscha) summer spawning migration in a tributary of Prince William Sound, Alaska.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate WPY2022 anadromous anadromy biological biosphere book chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water schooling semelparity semelparous spawning behavior spawning migration stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Chum Salmon spawning migration
Chum Salmon (Oncorhynchus keta) spawning migration in a tributary stream of Prince William Sound, Alaska. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Dolly Varden Dolly Varden Char Dolly Varden Charr Dolly Varden Charr southern form Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Salvelinus malma Salvelinus malma lordi Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water semelparity semelparous spawning behavior stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Coastal Cutthroat Trout in stained water
Coastal Cutthroat Trout (Oncorhynchus clarkii clarkii) schooled in tannin stained water. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Coastal Cutthroat Trout Copper River Delta Dolly Varden Dolly Varden Char Dolly Varden Charr Dolly Varden Charr southern form Ecology Infraclass Teleostei Neopterygii Oncorhynchus clarkii Oncorhynchus clarkii clarkii Pacific trout Phylum Chordata Salmonidae Salmoniformes Salmoninae Salvelinus malma Salvelinus malma lordi Seacam underwater housing South Central Alaska Spring Subclass Neopterygii Teleost Teleostei United States Vertebrate biological biosphere chordate diadromous diadromy fresh water freshwater in situ iteroparity iteroparous living lotic migratory naturalworld nature organism salmonography.com searun stream under fresh water underwater underwaterphoto wild wild salmon wild trout
Dolly Varden school
Dolly Varden (Salvelinus malma) schooled in pool. Aspect Ratio = 2:3.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Dolly Varden Dolly Varden Char Dolly Varden Charr Dolly Varden Charr southern form Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Salvelinus malma Salvelinus malma lordi Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate biological biosphere chordate courtship diadromous fresh water freshwater in situ iteroparity iteroparous living lotic migratory naturalworld nature organism salmonography.com spawning migration stream under fresh water underwater underwaterphoto wild wild char wild charr wild salmon wild trout
Male Coho Salmon with schooled Dolly Varden
A Male Coho Salmon (Oncorhynchus kisutch) with Dolly Varden (Salvelinus malma) schooled in a pool. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Dolly Varden Dolly Varden Char Dolly Varden Charr Dolly Varden Charr southern form Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Salvelinus malma Salvelinus malma lordi Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate WPY2022 anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ iteroparity iteroparous living lotic migratory naturalworld nature organism salmonography.com semelparity semelparous spawning migration stream under fresh water underwater underwaterphoto wild wild char wild charr wild salmon wild trout
Pink Salmon and Dolly Varden
Pink Salmon (Oncorhynchus gorbuscha) and Dolly Varden (Salvelinus malma) staging in a pool. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Dolly Varden Dolly Varden Char Dolly Varden Charr Dolly Varden Charr southern form Ecology Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Pacific salmon Phylum Chordata Pink Salmon Salmonidae Salmoniformes Salmoninae Salvelinus malma Salvelinus malma lordi Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate WPY2022 anadromous anadromy biological biosphere chordate diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism salmonography.com semelparity semelparous staging stream under fresh water underwater underwaterphoto wild wild salmon
Jack among other Coho Salmon
Coho Salmon (Oncorhynchus kisutch) jack or precocious male among other coho on their spawning migration in a tributary of Eyak Lake, Alaska. Aspect Ratio = 2:3.
Actinopterygii Alaska Alaska wild salmon Animal Autumn Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromous fish anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ jack jack salmon living lotic migratory naturalworld nature organism precocial male precocious male salmonography.com semelparity semelparous spawning migration stream under fresh water underwater underwaterphoto wild wild salmon
Maturing Chum and Pink salmon
Maturing Chum and Pink salmon (Oncorhynchus keta, O. gorbuscha) staging on their spawning migration in a tributary of Prince William Sound, Alaska. Aspect ratio = 1:2.5.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Summer Teleost Teleostei United States Vertebrate aggression anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water semelparity semelparous spawning behavior spawning migration staging stream threat posture under fresh water underwater underwater photography underwaterphoto wild wild salmon
Spawning migration
Chum Salmon (Oncorhynchus keta) and Dolly Varden (Salvelinus malma) holding in a pool during their spawning migration in a tributary stream of Prince William Sound, Alaska. Aspect ratio = 1:2.5.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Dolly Varden Dolly Varden Char Dolly Varden Charr Dolly Varden Charr southern form Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Salvelinus malma Salvelinus malma lordi Seacam underwater housing South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ iteroparity iteroparous living lotic migratory naturalworld nature organism photographed under water salmonography.com semelparity semelparous spawning behavior spawning migration stream under fresh water underwater underwater photography underwaterphoto wild wild char wild charr wild salmon wild trout
Chum and Pink salmon spawning migration
Chum and Pink salmon (Oncorhynchus keta and O. gorbuscha) spawning migration in a tributary stream of Prince William Sound, Alaska.Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Oncorhynchus nerka Pacific salmon Phylum Chordata Pink Salmon Red Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Sockeye Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water salmonography.com semelparity semelparous spawning behavior spawning migration stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Schooled Sockeye Salmon approaching spawning
Schooled Sockeye Salmon (Oncorhynchus nerka) arriving on spawning grounds. Male in background is gaping, which is a threat display. Aspect ratio = 2:3.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Ecology Infraclass Teleostei Neopterygii Oncorhynchus nerka Pacific salmon Phylum Chordata Red Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing Snell's window Sockeye Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism salmonography.com semelparity semelparous spawning migration staging stream under fresh water underwater underwaterphoto wild wild salmon
Male Coho Salmon gaping during spawning migration
Potential alpha male Coho Salmon (Oncorhynchus kisutch) gaping during spawning migration. Aspect Ratio 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Class Actinopterygii Coho Salmon Ecology Infraclass Teleostei Neopterygii Oncorhynchus kisutch Pacific salmon Phylum Chordata Salmonidae Salmoniformes Salmoninae Seacam underwater housing Silver Salmon South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate aggression anadromous anadromous fish anadromy biological biosphere chordate diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism salmonography.com semelparity semelparous stream threat posture under fresh water underwater underwaterphoto wild wild salmon
Pink salmon spawning migration
Pink Salmon (Oncorhynchus gorbuscha) spawning migration in a tributary of Prince William Sound, Alaska. Aspect Ratio = 1:2.5.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Hartney Creek Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature near Cordova organism photographed under water run timing salmonography.com semelparity semelparous spawning behavior stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Chum and Pink salmon spawning migration
Chum and Pink salmon (Oncorhynchus keta, O. gorbuscha) spawning migration in a tributary of Prince William Sound, Alaska. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Hartney Creek Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature near Cordova organism photographed under water run timing salmonography.com semelparity semelparous spawning behavior spawning migration stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Chum Salmon spawning migration
Chum Salmon (Oncorhynchus keta) spawning migration. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Infraclass Teleostei Neopterygii Oncorhynchus keta Pacific salmon Phylum Chordata Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature organism photographed under water salmonography.com semelparity semelparous spawning behavior spawning migration stream under fresh water underwater underwater photography underwaterphoto wild wild salmon
Chum Salmon spawning migration
Chum Salmon (Oncorhynchus keta) upstream spawning migration. Aspect Ratio = 1:2.
Actinopterygii Alaska Alaska wild salmon Animal Biology Chordata Chum Salmon Class Actinopterygii Dog Salmon Ecology Fish Hartney Creek Humpback Salmon Infraclass Teleostei Neopterygii Oncorhynchus gorbuscha Oncorhynchus keta Pacific salmon Phylum Chordata Pink Salmon Reproductive behavior Salmonidae Salmoniformes Salmoninae Seacam underwater housing South Central Alaska Subclass Neopterygii Teleost Teleostei United States Vertebrate anadromous anadromy biological biosphere chordate courtship covering redd diadromous diadromy fresh water freshwater in situ living lotic migratory naturalworld nature near Cordova organism photographed under water run timing semelparity semelparous spawning behavior stream under fresh water underwater underwater photography underwaterphoto wild wild salmon